Gorgonops, I'm sure you can set me straight on this, as much of what I'm saying about the electrical compatibility of the Twiggy vs. the Sony drive is based on my peripheral knowledge of planned Fileware implementation...
So... here's what I *believe* would count as "corrections"... I've referenced some documents like the
Lisa FAQ at sunder.net, "Inside Macintosh", and an old IWM data sheet among others, but some of this is going from memory so it may not be perfect.
1: The reference to "widget" driver was discussed a while ago in one of our long threads about alternate ways to add mass storage to an original Macintosh. It's a red herring. That driver is in the "soft" version of the 64K Mac ROM that was loaded on Lisa 2s/"Macintosh XLs" when they were emulating a Mac. It's not in the "real" Mac ROM.
2: "Parallel vs. Serial": The IWM chip was designed with the Fileware drive in mind (the part predates 3.5 inch floppy drives by several years) and was used in the Lisa. (The IWM supports two modes of operation for the "control lines" present on the 19 pin external connector; there's the "Mac Way" where the lines are used to communicate semi-intelligently with the control registers on a Mac drive, and it also supports using the lines to generate raw stepper-motor control signals compatible with a Disk ][-style drive. If Fileware for the Apple II/Apple /// had ever become a shipping product I imagine both modes would be available on the associated controller.) The main difference between the Lisa and the Macintosh is in the Lisa drive I/O was handled by an embedded 2Mhz 6504-based computer not the main CPU, but that computer lived on the I/O board in the main card cage and not on the drives. The drives themselves are still "dumb"(-ish), like the Mac's drive. It was the Lisa's Widget and Profile hard drives that connected via parallel-(ish) ports.
There is one interesting thing, however: There were two versions of the Lisa 2, the 2/5 and the 2/10 (when you converted an original Twiggy Lisa to a Lisa 2 it became a 2/5), and the two systems had different I/O boards.
This is covered in a section of the Lisa FAQ. In order to work with the 20 pin Mac-style floppy drive which was used after the conversion Lisa 2/5s (or former Lisa 1s) required a "Lite Adapter" be present in the card cage assembly to covert the *26 pin* Lisa Twiggy floppy connector. The adapter *mostly* remaps the pins but in addition it generates a 3.5-inch compatible PWM speed control signal in place of the Twiggy-appropriate signal. I had noticed from photos/video footage of a Lisa 1 that the drive cables *did* appear to be wider than a Mac's drive connector cable, and this is why....
HOWEVER, if you look at the photos of the Twiggy mechanism inside the prototype Mac it appears that is only has 20 wires, and the circuit board on the drive appears to be different compared to the Lisa version of the mechanism. So... this seems to be solid evidence that this prototype is using genuine intended-for-the-Macintosh parts rather than being fraudulently thrown together using a Lisa 1 Fileware drive and a Twiggy faceplate. Whether the use of the 20 pin connector indicates that the drive is directly compatible with the 3.5 inch drive is another question; the PWM signals are obviously incompatible between a *Lisa* Twiggy and a Sony drive, but since the Mac Twiggy uses a different circuit board all bets are off. Even if the two are electrically compatible if the board is fitted with the wrong ROM for the Twiggy drive (or the ROM is for the Twiggy drive and an externally-connected 3.5 inch drive is connected, for that matter) the drive that doesn't match the ROM might be spinning at the wrong speed to work properly even if you had an appropriately formatted system disk inserted in it.
More interesting to me now is that the floppy drive icon is indeed the 3.5" one, which guarantees that a Sony driver is in ROM. But as you point out, possibly not one compatible with the final drive considering what we know about the minor update during its production run.
So, I've been thinking about the floppy drive icon question and one possibility has occurred to me:
There undoubtedly was a transition period in which both Twiggy and Sony Macs were kicking around the labs. The Mac-adapted Sony drives were probably in short supply for at least a month or two, during which it would of been impractical to suspend all other work on the OS while the bugs were being worked out of them. One of the aspects of the Mac OS which is very clear from looking at the documentation (and from reading Folklore.org) is that the RAM-loaded part of the OS and the ROM are very tightly intertwined, and until the "gold release" date both were being updated on a regular basis as bugs fixed in one part required a change in the other part. (And as a result, changes that touched the ROMs would require all the software developers to immediately install a fresh set of EPROMS in their development machines.) The floppy driver in the Mac ROM is fairly self-contained and for normal use (IE, not formatting a disk or something) abstracts a floppy into appearing as a linear block device. (The vMac and BasiliskII developers leverage this property by patching out the floppy driver and replacing it with a disk image driver for emulation purposes.) What if for a certain period of time there were *two sets* of development ROMs being passed around the labs, one for Sony prototypes and one for Twiggys, and at a certain point the "blinking disk" icon was replaced with a picture of the Sony disk but disk driver remained Twiggy compatible for the developer machines that needed it? That would nicely explain why this prototype can't boot from an externally-connected 3.5 inch drive. It *may* be able to boot from the Twiggy, but generating the disk for it may be well-neigh impossible. A dump of its ROM would probably reveal the truth if someone knowledgeable enough were to isolate the disk driver and compare it with the shipping version.
(I suppose in principle if one *were* able to extract a Twiggy-driver from the ROM of this system one could go about generating a boot disk for it by connecting the Twiggy mechanism to something like a Mac Plus with a hard disk, loading a custom init that replaces the ROM disk driver with the Twiggy version, and using a custom formatter program to generate an appropriate disk to which a pre-release System version that might be compatible with the rest of the ROM could be copied to.)
I know Apple doesn't give away ice in the winter, but why give a non-functional computer as a gift? :
The computer was not a *gift*. The story is that it was sent to a sculptor to use as a model for a medal that was going to be given to members of the Mac development team. Supposedly the sculptor finished the job but Apple never requested he send the computer back to them so he kept it. Assuming the story is true your idea that this machine was "assembled from protoparts" specifically for this task doesn't make any sense; why would they go through the trouble to "assemble" a computer that doesn't even need to work when they could just grab one, regardless of condition, that looks enough like a Mac to serve the purpose? (If they were actually wasting the time putting something together why not just send him an empty chassis with case plastics screwed to it?) It makes a lot more sense that they either snatched this off someone's desk (to be replaced with a new Sony-equipped unit) or out of a storage room/loading dock/garbage bin where obsolete Twiggy units from internal developers /shipped back from places like Microsoft were piling up. For all we know this machine could be the very one sitting under a stack of papers in one of those pictures linked earlier, rendered obsolete by a Sony-drive model shipped to Redmond the previous week.