I've pulled tops off of several brands of hard drive, to varying degrees of success.
Quantums are really easy, because they're (usually) the toilet-bowl design with the stamped metal top that easily screws down, with no odd protrusions into the lower bit of the case. Most of them work fine with or without their tops, but some single-platter drives won't operate properly exposed because of the peculiar implementation of the AirLock system.
Conners can be problematic, as it's easy to hit the platters when removing the bowled top casing. Other than that, most of the ones I've run exposed have exhibited no problems. In fact, several have run without their tops when they wouldn't run at all otherwise.
Seagates are a mixed bag: some work fine, some don't. Medalists are usually fine, but (sub-10GB) Hawks and Barracudas typically freak out and won't even spin up if the top is off. U Series drives usually work nudified, but they're difficult to disassemble with both the sealing tape and the shock-absorbing "condom".
WDs work fine open, but older ones are almost always sealed with tape, which is close to impossible to remove in one piece. When I reseal the taped drives (of any brand), I use thin strips of duct tape.
IBMs are also very cooperative, even the notebook models. A couple of odd UltraStars are the exception, because their casing is split down the middle.
Fujitsus, Maxtors, MiniScribes, and Toshibas typically run exposed well with no real issues.
As for positioning systems, there have been many different approaches over the years. Early drives often just used stepper motors, counting the steps the motor moved the heads to tell where on the disks they were positioned.
Some later drives, Quantum and Sony for sure, used an optical tracking system. These drives employed a small attachment to the armature, a small piece of glass or plastic with a bar code-like grid on it, read by an infrared LED and pickup. The electronics would count the movement of the gradients on the tracking grid, and thus keep track of where the heads were positioned.
As a side note, on Quantum ProDrive LPS drives (from 40 to 230MB, employed quite frequently in Macs), the little "tick-ticka-tick-ticka-tick" sound emitted every 15 to 30 seconds is the sound of the electronics recalibrating the heads, just in case it missed counting a gradient.
Then, modern drives use embedded servo information. Some drives would dedicate an entire platter to maintaining servo information, where others embed the servo data alongside the usable data.
I've never tried swapping platters or head assemblies, though, as most of my problems have been easily solvable (stiction, damaged PCBs) or unsolvable (extreme shock or water intrusion).
As for swapping PCBs, older drives (<4GB) are picky about firmware revisions and all, but modern drives work just fine so long as the boards are from the same model and capacity.