[NITPICK]
1) The 7500, 8500, 7600, 8600, and 7300 motherboards have the same form factor, but the 7600/8600 changed the power supply connector. The 7300 uses the 7600 style connector.
2) There were two version of each of the 8600 and the 9600. There was the original 8600/9600 with the 343S0280 - 343S0283 ROM chips and cache on the 9600 logic board. These original models use the same CPU cards as the X500 machines.
Then there were the 8600 Enhanced and the 9600 Enhanced, which are also called Kansas machines. They use the CPU cards witht he Mach V PPC604ev CPUs. They have 343S0380 - 343S0283 ROM chips. The 9600 Enhanced has no cache chips on the logic board but otherwise (except for minor rearrangement of CPU slot pins) is identical to the earlier 9600 logic board. I have never mapped it out, but I suspect the only real difference in the CPU slot pinout is more 3.3V supply pins on the Mach V cards.
3) ROM for the 8500 and 7500 and 7600 (and 9500 and most 7200s) were identical. So moving a ROM DIMM from an 8500 to a 7500 or 7600 would not change anything. Moving said ROM DIMM to a 7300 would make a change, as the 7300 uses the same ROM as the original (non-Kansas) 8600/9600.
[\NITPICK]
I suspect what the OP has there is a 7300 variant that actually supported the Mach V processor (changes in the CPU slot pinout). The 7300 came out with the 8600/9600 Enhanced, IIRC, yet had the ROM of the original 8600/9600 and did not support Mach V.
It would have made sense for there to be an unreleased 7300 which was consistent with the 8600/9600 Enhanced and had a Kansas motherboard supporting Mach V CPU cards.